Rachel wakes after her surgery to get a new eye courtesy of clone-Krystal. She's in an all-wooden room and appears to be alone, but she finds an envelope on the nearby desk. Removing her bandage, she finds one impressive, slightly-cyber-looking eye.

Sarah, Felix, and Mrs S bring Kendall Malone to Scott, who will be doing tests on her and hopefully keeping her hidden from Castor. We also learn that the reason she looks different from Sarah is because the two strains of DNA combining cause her traits to manifest differently.

Dr. Coady tells Rudy that they'll need to go after the original but the only clone that hasn't gone underground is Alison, who is still busy campaigning. But Helena is also still with Alison and Donnie, who surprises Helena with Jesse, the tow-truck driver whom Helena hit it off with last season.

Cosima goes to visit Shay to apologise for accusing her of shady dealings and Shay is not happy because of the threats Delphine laid on her. Cosima is thoroughly apologetic but can't let Shay in on all the secrets of Clone Club so it looks like this relationship is kaput.

Ferdinand pays a visit to Delphine at Leda. He wants to run off with Rachel and be in charge of Top Side and Leda. But Ferdinand's in for a surprise, as Sarah is there to offer a one-time deal. She also tells him that she was the one who was pretending to be Rachel the last time he came to visit. Sarah is willing to offer up samples of Malone in exchange for destroying Castor and Coady. Delphine says he's the lesser of two evils.

Cosima is the one brought in to take Kendall Malone's samples. Though Kendall's snappy with the biting comments, Cosima's warmth and compassion convinces Kendall to give up the samples to help save her life.

Sarah and Felix track down Mark and Gracie to a seedy motel room, and Felix even kicks the door in to get to them. Mark seems willing to help without much work on Sarah and Felix's part. He had thought he'd spend his last few months with Gracie before he died but this is his only chance for a cure.

Mrs. S tells her mother that if need be, she'll destroy her genetic material with an acid bath. She also wants revenge for John. She promises that if anything goes wrong, she'll be putting Kendall in that acid bath.

Delphine wants Rachel moved to a secure facility and orders Nealon to have her moved. After Nealon leaves, Delphine notices that "Rachel" has unusually manicured nails and then "Rachel" wakes up and smacks her in the face. She determines quickly that the patient in the bed is Krystal, who thankfully still has her eye. She wants to know what's going on and why weird things keep happening to her. Did they just need to take a sample of her eye? We're certainly glad she still has it.

Rachel is also confused about where she is too. She's being watched though security cameras and sees the name "Finch" nearby. She demands to know who's keeping her there but gets no response.

Helena catches Jess up on everything that happened since they were away, including the prison break and soap-making. She's all prepared to hook up with Jesse in his truck but then she gets a call from brother-sestra Felix and learns that her family needs help. Jesse's willing to wait, though.

Alison is at the polls with Donnie, and Rudy is lurking nearby to follow her and her campaign school-bus. Donnie knows they're being watched though and gives Alison a heads-up and texts Sarah to let her know. Sarah is also tasked with punching Mark in the face repeatedly for a cover story, we think.

Alison's campaign bus makes a pit-stop to lure Rudy out of his truck. He heads into the garage where Helena is waiting for him. She's taping weapons to her hands, saying it's "prison rules." He doesn't want to tape his knife so she kicks the duct tape into his nose. Some time later, a bloody-faced Rudy calls Coady to say he's found the original, turning the phone to show Kendall, and sends Coady the location. But it's actually Mark in disguise.

Helena still has Rudy to fight, though with his glitches it's not much of a fight. She gets a screwdriver into his arm, forcing him to drop his knife. Cody arrives at the location and sends some of her goons inside. But then Ferdinand shows up and shoots here driver dead and apprehends Coady, saying she won't find what she's looking for there.

A dying Rudy shares childhood memories with Helena. Rudy ponders their similarities, but she tells him they're not alike, given that Rudy is a rapist. But she does lovingly stroke her dying brother's face.

Mrs. S gets Kendall to share more information. Duncan had visited her some years after his original harvesting to admit what he had done and the fear he held for his life. Everything had gone wrong but he was able to hide the original, Kendall, and there was one child outside of it all that was hidden in the foster system (Sarah). She also mentioned something about Neolution. Kendall put Sarah in Mrs. S' custody because she had the connections to keep her hidden; it was the one last piece of her own foul life. Kendall and Mrs. S are able to have a moment of reconciliation.

Delphine has Dr. Nealon in holding. She wants answers on who he works with and Rachel's whereabouts. He tells her to turn on the screen, which she does, and finds the surveillance feed of Rachel. Delphine also seems to know about Neolution, something Nealon claims has infiltrated and manipulated all the moving pieces of this, including Leda, Top Side, and Castor.

Ferdinand comes to collect samples from Kendall, while it's implied that Ferdinand's man taking the samples is actually part of Neolution. Nealon offers Delphine a place in Neolution but when she refuses, he attacks her and displays a worm coming from out of his mouth. Delphine gets a shot off into Nealon's chest, who ominously tells her that she won't live till morning. She takes the worm out of Nealon's mouth using a pair of pliers and calls Sarah to tell her that she can't let Ferdinand have the samples and that both Leda and Castor are run by Neolution. She says to tell Ferdinand that Rachel is still alive and was taken by Neolution.

When Ferdinand hears this, he goes on a rage and beats his goon with a bat. The man admits to being a Neolutionist, and Ferdinand hates Neolutionists so he's now apparently part of Team Leda-Clone because he refuses to hand over Kendall's samples. He wants Kendall hidden, Mrs. S has connections, and tells them all to head out while he'll dispose of this Neolutionist's body.

Delphine goes to Shay's to apologise to her and gives them their blessing, promising to not stand between her and Cosima. She also tells Shay that she can tell Cosima everything if she wants. Looks like Shay has more secrets after all. She also tells Shay to give Cosima a card; it's of the Dyad Institute and has 324B21 written on the back.

The gang all comes together for a dinner party at Bubbles, where Alison gets the call that she beat Marcie Coates. The only one not present is Kendall, who is resting up before the trip she'll be taking to disappear. Sarah and Art note how not everyone is present, and everyone toasts to Beth. Helena also toasts to Kira's "little face." Alison then toasts her victory and that they'll all face whatever comes next together. She also thanks Donnie for standing by her through everything. Cosima then gets called outside by Delphine, who asks her to keep the genome sequence and Kendall secret and safe. The two mend fences and Delphine kisses her goodbye. Delphine then heads to her car in the garage and upon realising she's not alone, turns to ask, "What will happen to her?" She is then shot in the chest. The identity of her shooter is not shown.

Rachel finally gets a visit; it's a little Leda clone named Charlotte who says that "she" said that Rachel will be her new mum. The "she" in question is Professor Duncan, Rachel's mother, who steps out to identify herself.

Meanwhile, Mrs. S, and Kendall arrive in Iceland where Sarah is reunited with Kira.

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Sarah, Felix, and Mrs. S have made it to London safely in search of the original source of the genome.

Rudy's still glitching and his condition is only worsening. Dr. Coady assures him that their men will track down the original genome in time to save him.

Helena spends some quality time with Alison's daughter, who is preparing for a karate match. Helena teaches her some not-so-ethical moves, namely eye-gouging, but Gracie interrupts to point out that Helena needs to not do things like that anymore. Helena agrees, saying she's a mother now and has turned over a new leaf. Gracie then goes to ask Alison for the day off to head to a doctor's appointment. Alison's looking for Donnie, who didn't come home because he was knocked out cold at Bubbles after his fight with Jason.

Cosima looks through Shay's folder and is being distant from her after learning the truth of her background and possible involvement with Castor. She seems unsure how to proceed just yet.

Donnie gets a visit at Bubbles from one of Jason's men to say that he's been taken out of the drug-dealing machine because Jason no longer vouches for him. Pouchy wants the pills back and the man takes the nitrogen case ransom, except Donnie has no idea what it is since Helena was the one who hid it there in secret.

Mrs. S has gotten in touch with some of her old contacts in London to sniff out information on the original clone. But her old friend Terrence ends up brutalised at the hands of one of Ferdinand's goons. Terrence had been working with Ferdinand and was supposed to let him know if/when Mrs. S ever came back to town, though it was entirely clear that his allegiances were truly with Mrs. S. Following a rousing round of karaoke, Mrs. S heads over to where Terrence had said to come meet him, where he was in the midst of being tortured for information on what S was looking for.

Helena catches Donnie when he's looking for the pills to return. She insists on patching him up and for answers on the source of his injuries. Helena learns the truth about the soap-making being a cover for dealing drugs but she is far more upset to learn that her freeze-dried babies in the tank have been stolen. She's ready to go hunt them down but Donnie tells her not to, and that he'll take care of things for the family which he says she is also a part of. Helena seems touched by the gesture.

Cosima goes to Delphine for help at Dyad. She says how Sarah, Mrs. S, and Felix are in London in search for the Castor original and that help and resources will be needed to stop them from beating Castor who is hot on their trail.

The London trio find a mortally wounded Terrence who gives S enough information to scare her off the trail but not Sarah, who swipes Terrence's phone and calls his last contact to try to follow the lead. Felix insists on coming along to make sure she's safe, despite not approving of her choices.

Donnie goes to return the pills and is hoping to get his money back too but instead, Pouchy says that they're working for him now. Helena had come inside to pretend she was Alison and get the tank back safely in her hands. But as the two are walking out, one of the dealers threatens Alison and Donnie's children and Helena's response is to shove Donnie out of the warehouse with the tank safe in his hands while keeping herself locked in there to dispose of them. She later exits the building with a giant machete, soaked in blood, and carrying a giant bag of cash. She's gotten Donnie's refund and thanks him for helping to save her babies. She needs to shower up and Alison is shocked to return and find a battered husband with a newly packed freezer full of cash.

Delphine goes to Shay's place to threaten her into admitting her involvement with Castor. She threatens to kill her and make it look like a suicide but Cosima calls to let her know that it wasn't Shay after Gracie calls her to admit that she had ratted them out to Castor after they promised that she could be with Mark, whom she still loves even after processing the truth about his being a clone.

Sarah and Felix follow the leads to a prisoner numbered H46239 and named Kendall Malone who is the original genome. Ferdinand calls Delphine to say that he's learned about Sarah not actually being in her custody. So now the playing field has been evened since she had previously muscled him out because of his insidious scheming done with Rachel. Sarah has broken into Kendall Malone's flat. Unexpectedly, Kendall Malone is an elderly woman who is plenty tough. They end up facing off with another but Mrs. S arrives and surprise! Mrs. S is the daughter of Kendall Malone! She had been tested on while in prison by Professor Duncan and Malone carries the genetic DNA of both Castor and Leda because she had absorbed a twin when she was in utero. But outside, Felix is starting to panic as he thinks he's spotted someone (probably Topside) that's going to come in and grab them. Mrs. S is prepared to burn down her mother's apartment and kill her mother to destroy the Castor DNA but Sarah implores her not to, and to understand that they need her since she is both Castor and Leda.

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Delphine goes to get her nails done and the woman doing her nails is a clone we've not met yet, save for some footage of her about to hook up with Rudy. She shares the story of this crazy hookup with Delphine. Turns out Krystal's boyfriend Hector ended up dead following this encounter with the "twins." Delphine also has some not-so-nice plans for Rachel, namely wanting her to be eliminated.

Sarah open to Cosima about how much she's been thinking about Beth and wants to finish what she started. She doesn't want to lose another one of her sisters. Cosima assures her that no one has been lost yet. Also, Cosima has been staying at Shay's place, as the two are getting cozy.

Felix brings Helena over to Alison and Donnie's, since it's their turn to look after the "crazy" one. Alison thinks Helena can help them to make soap as part of their newly-acquired Bubbles business. Helena's also good with kids so she gets on quite well with Alison and Donnie's children. She's also secretly lugging around her nitrogen tank that she believes is carrying her babies.

Sarah speaks with Scott and realises she needs to speak to Rachel, so Scott helps set up a Skype call. Rachel's on to the fact that Delphine wants her gone. She offers to translate the book in exchange for them getting her out of Dyad and says she wants to leave a quiet life in Taiwan. She then offers up the name of clone-Krystal and says she wants them to help her steal the identity so she can get out. As proof of her knowledge of the book, she has translated the copy of the page of the book that Scott offered up. The message reads, "In London town we all fell down and Castor woke from slumber." Mrs. S is skeptical of doing this plan and argues with Sarah over how to proceed. Gracie overhears the argument and the group assures her that Mark is different from the rest of the Castor clones.

Cosima brings Shay to Dyad to show her around but Delphine breaks up the party with bad news regarding the ongoing research of the sequence. There was a test trial on another ill Clone but it didn't work and she died. Delphine wants Cosima to have another stem-cell treatment from Kira but Cosima refuses to harvest her like that again.

Donnie and Alison bring Helena to Bubbles and have her begin working on some soap in the back. Helena playfully flirts with Donnie by comparing him to a baby ox. But Donnie has bigger concerns on his mind, confronting Alison about his displeasure at Jason having kissed whom he thought was Alison. But this conversation will have to wait, as Gracie arrives to also begin working at the shop. Also Helena is hiding her nitrogen tank in the back. Is that safe? The first reunion between Helena and Gracie is not so tense the way you might have guessed. Gracie apologises for her miscarriage and Helena says it's okay and that Gracie will be an auntie to her baby.

Scott returns home to find Rudy lying in wait with Scott's cat on his lap. As it turns out, someone has tipped Castor off about Duncan's Dr. Moreau book and the hidden code inside. How did he find out about that?

Krystal is blabbing her story once again, this time to Felix who has come in with a fake North American accent to get his nails done. He's basically flirting with her and it's hilarious. He's also pumping information out of her so Sarah can document it so her identity can be stolen. Krystal also expresses her concerns about the strange happenings that have been going on in her life, including how her boyfriends are basically on a two-year rotation to the day. Felix wants to tell her about Clone Club but Sarah refuses to let him, and Felix needs to hurry up and go back to steal her wallet before heading out. Felix also finds a notebook indicating that Krystal has been more thorough in her investigation of the things going on in her life, even documenting them in a little notebook. At least dear Felix was nice enough to leave her with some words of encouragement.

Alison confronts Jason on the kiss but he ends up kissing her again. He's all but offered himself up on a silver platter to be Alison's plaything but Alison is insistent that she is married and he can't do anything. Donnie then texts Jason from Alison's phone to invite him to Bubbles and the two get into a slap-turned-punch fight that leaves Donnie knocked out cold in the work room.

Cosima and Delphine are having a moment in the lab but they're interrupted by a call from Scott, who tells Delphine about the book since it's been stolen by Rudy at the threat of harm to Scott's cat. Delphine is furious at not having been told about the book and Cosima tells her that the book is gone. In actuality, they did make a copy of it but they're not about to tell Delphine that any time soon. The thing is, Rachel was the only other person who knew about the book so she had to have told a mole who tipped off Castor. Delphine interrogates Rachel, who isn't offering up an explanation over how she got the information out.

Cosima, Scott, and the rest of the team is working together to help Rachel make a break for it. Cosima brings Delphine her resignation from Leda. Rachel makes it to Mrs. S' place where the rest of the group are waiting. They give Rachel all of Krystal's stolen IDs and sit Rachel down so she can translate the entire book from start to finish, after which she'll be free to go. Meanwhile, Cosima admits that she had a near-death experience and the only thing that allowed her to come back from it was a vision of Delphine. She tells Delphine she needs to let her go but the two end up kissing before Delphine says that she should have trusted her.

As soon as Rachel has finished translating the book, Dr. Nealon and other Leda team members come in. Rachel is rushed back to Leda after having some sort of episode and Leda gets their hand on the book. Nolan claims that Rachel may never recover but in actuality, he and Rachel were in on it together. They had Krystal kidnapped so Rachel can have her eye and head off to London, where the original genome is located. Mrs. S has also figured out the message of the code so now they, along with Sarah and Felix, must head off to London too. Delphine is furious to not have been told about the copy of the book so Cosima's resignation has been accepted, but not before she hands her a file on Shay, who appears to be the Castor mole.

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Mrs. S is able to track down a freshly-escaped Sarah and Helena. Helena, now knowing that Mrs. S was responsible for putting her in Castor's captivity, wants to kill her but Sarah stops her from doing so... for the time being anyway. Helena wants nothing more than to kill her. Helena had also been shown to be comforting Sarah regarding Paul's death, saying that he was a good man in the end. Sarah is emotionally exhausted and doesn't want to discuss it.

Alison and Donnie go meet Connie, Alison's mother, who owns the Bubbles soap shop that they're hoping to take over in order to use as a storefront for their pill-pushing. Alison seems to be insecure around her mother and appears to be feeling inadequate. Connie has even put up a campaign sign up supporting Marci, Alison's rival. Connie is stalling on signing over Bubbles to Alison and Donnie. Alison later has a campaign event and Felix attends to help Alison with it all.

Cosima is enjoying her post-hookup bliss with Shay but they're interrupted by Delphine, who wants a urine sample from Cosima because she says her numbers are worrying. Shay eavesdrops on their conversation from the bathroom. Cosima texts Aliso to ask her for a urine sample, after having a conversation with Scott about trying to get Rachel to decode the book. Alison is interrupted right before the event by a call from her mother claiming she's having a heart attack.

Donnie goes with Alison's ex Jason in order to drop off a payment to a higher-up drug dealer but when they arrive, they realise Donnie has the wrong envelope. The money is with Alison and Donnie instead has an envelope full of campaign buttons. Jason is sent to retrieve the money while Donnie must stay in the captivity of the drug-dealers.

The fight between Helena and Mrs. S hits a breaking point. Helena repeatedly hits her, insisting that Mrs. S hits her back. Mrs. S only lands one punch and then hugs her, saying repeatedly how sorry she is. A struggling Helena eventually relents and appears to make peace (somewhat) with the situation.

Alison arrives at her mother's store o find that her mother is fine and only had a panic attack of sorts. Connie then says that she can't sell Alison the store and Alison keeps getting texts from Felix who is saying that she's about to miss the photo-ops. Alison delivers an impassioned speech to her mother for never deeming anyone good enough. When Cosima arrives at the campaign space to try to get Alison's pee, she is mistaken for Alison and must now pretend to be her for the photo-ops. Felix gets her looking the part and pushes her to go through with this facade because if she wants help from Alison, then she needs to return the favour.

Back at the lab, Rachel is brought in for testing by Delphine and Scott reveals to her without Delphine seeing that he has her father's book. She claims to want to be taught the boardgame that Scott and his friends had been playing and Delphine gives permission, not knowing it's a rouse to allow them to be able to be alone in the future.

Alison arrives back at the campaign centre and spots Cosima upstairs doing the photo-ops and must hide while also being met by Jason, who tells her about what has happened regarding Donnie being held captive and the money. Alison texts Felix to help her retrieve the envelope from the table but then her mother arrives. Alison pulls her away to keep out of sight and then tells her that Alison was conceived in-vitro with the use of sperm that wasn't from Alison's father. She only ever wanted the best for her. Alison runs into Jason in the hall, and asks him to keep her mother preoccupied while she goes off and retrieves the envelope. Meanwhile, Felix is telling Cosima that she should hide in the loo until he can get Alison to come in and give her a urine sample.

Alison tracks down Felix and tells him about the envelope mix-up and has to admit that she and Donnie are drug-dealers and now Donnie is being held captive by a drug-dealer named Pouchy who coincidentally is the same man who cut off Vic's finger.

Jason is able to sweet-talk Alison's mother, seemingly making her believe in Alison more and in turn make her more compliant to sign over the business. Alison is able to swap out the envelopes and meet Pouchy's niece in the parking lot and is forced to sit there while the money is counted when she's supposed to be inside making her speech. So poor Cosima must go in and make the speech on behalf of Alison. Pouchy gets the call that the money is in play just before his muscle is about to cut off Donnie's nose.

Cosima is struggling to make a speech up and is coughing so she uses that as a guise to go backstage where Alison has made it back in and is able to swap back in to getting onstage and making her speech. Instead of the meticulously prepared one, she is inspired at seeing her mother in the audience to swift her tactics and say that her campaign is about family. The speech is well-received by the audience and Marci looks worried.

When Cosima heads back to go to the bathroom she is approached by Jason who believes her to be Alison and plants a kiss on her. After the speech, Alison learns from Cosima about the kiss from Jason and she seems quite surprised but perhaps... flattered? Alison then decides against giving Cosima her pee and says they need to be honest with Dyad because they need their help regarding their health. Cosima then goes hiding in the stall because Alison's mother has come in. Connie congratulates her on her speech, saying that she's earned her vote and gives her the form to sign over Bubbles. She also insults Donnie in the process, saying that she knows that when the time is right she will learn that she deserves better and insinuates that perhaps she can get back together with Jason. Alison then decides to spite her mother and reveal Cosima to her and the fact that she is a clone. She says that she didn't fool Alison's father the way that she had thought and instead she was implanted with a clone. Cosima leaves, feeling awkward. Connie doesn't believe Alison, thinking that Cosima must be her half-sister instead and that even though the donor was supposed to be exclusive, perhaps Cosima's mother received a sample too. She also had initially thought that Cosima was mixed-race because of her appropriative hairstyle.

Sarah learns that Mrs. S has patched things up with Helena. They fill each other in on what they've missed, including Paul's death and Gracie now staying with Mrs S.

Cosima opens up to Shay, saying she has a pretty serious health issue. Rachel is shown to be playing the farm game with Scott and she wants the book but he tells her that the book does not belong to her, nor was it left for her. Cosima is shown to be bleeding out of her vagina and Rachel begins to decode a copied page from the book given to her by Scott but says she'll only give the information to Sarah.

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Sarah has a dream in which Kira appears on the base and leads her to a room where Sarah can see herself lying on a slab and having one of the male clones giving a blood transfusion of his own blood into her body. It's unclear as to whether this is some sort of recovered memory or simply a manifestation of her fears. The real Kira is of course still in Iceland and Skypes with Mrs. S and Felix. Mrs. S is still using her connections to try to find Sarah. Gracie brings them a tray of tea, saying it helps her to have something to do.

Paul meets with another agent and tells him about Doctor Coady's unsanctioned research but Paul needs to give more proof if actions are to be taken against her. Paul heads to the base and finds that Sarah is really sick. He gives her some water and she pleads with him to help her escape and he says he's trying to help her out of there. When Dr. Coady comes in, Sarah claims that she did something to her to cause her sickness, despite Coady's claims that she's trying to help Sarah and gave her an anti-inflammatory. Sarah enquires after Helena and Coady says that if the desert doesn't get her, then Rudy will. Rudy is driving a patrol through the desert, and a hungry Helena proceeds to eat one of her scorpion friends.

Cosima is enjoying the post-hookup bliss with Shay, who asks about Sarah, claiming that Cosima said her name in her sleep. Yeah. Sure. But when Cosima heads into work, she is met by Delphine who is waiting for her. She knows that they got their hands on a Castor-brain. She also points out that there is a protein present in Gracie's brain that was also in the Castor-brain, which clearly warrants a follow-up. Gracie learns that she is now infertile, and she feels relieved. As Cosima is leaving, Delphine says she misses her but Cosima doesn't look too pleased to hear that.

Alison and Donnie are enjoying their new funds from their drug-selling scheme. They proceed to make it rain in their room and get walked in on by one of their children who is promptly ushered out. She later meets with Jason, her drug-dealing ex, to pay back the money she owed on the supply. She wants to take her drug-dealing to the next level but he doesn't think she's ready for that, even more so when Donnie rolls up in a fancy new car and Alison chides him for coming and being ostentatious in his spending. Alison needs a business for a front through which to launder the drug-money. Alison proposes that she and Donnie take over "Bubbles," her mother's store. Alison has Jason come for a visit, despite Donnie's being disinclined to want to bring him along.

Paul asks another soldier on the base about what's going on and the soldier says how things have changed since Paul left. Rudy had brought a sick girl to base, whom Coady treated. But after that, things become more secretive and the boys were ordered to record their sexual activity and take samples. Everything is in Coady's office, which is restricted. He then speaks with Mark, who says how Sarah helped retrieve the Castor DNA and that he brought her to the base because he was with Rudy who would have otherwise killed her. He also admits that he fell in love with Gracie, once Paul pushes him to speak freely. Mark helps him retrieve the keys to get into Coady's office and get more proof. Mark is supposed to stand watch but he comes into the office and looks through the research with Paul, and he tells Mark that the defect is contagious through sexual contact. Paul then places Coady under arrest for her crimes and she admits to infecting Sarah with the Castor pathogen. Sarah hallucinates meeting Beth, who shares the story of how she met Paul. Sarah is wracked with guilt over not being able to save her sisters and Beth speaks about how being consumed by all of the clone-craziness. Beth tells her to stop asking "why," but start asking "who."

Felix asks Scott for help in the form of showing him where Rachel is. This conversation takes place when Felix brings Gracie in to Leda for more testing. He threatens and humiliates Rachel, demanding to know where Sarah is. But she doesn't want to help, only wanting to get out. Felix leaves in a huff.

Paul calls in his arrest of Coady and shares his findings, including the fact that she has been sterilising women. He is told that it will take six hours to get an extraction team and he must hang tight and manage things until then. He is also advised to leave Rudy out in the desert. Rudy gets a call from someone back at the base, learns of the situation, and proceeds to head back.

Delphine is shown to be monitoring Cosima's new relationship with Shay. Cosima goes to make out with Shay but gets a call from Scott who says that based on the paintings Rachel has been doing, he thinks she may know the code in her father's book.

Paul visits Sarah and tells her about the sterilisation of women and also that Coady has stated that Sarah hasn't been showing any of the same symptoms. Coady seemed to know that Sarah would beat it and Sarah demands to see her. The clones have the same defect but while the boys' brains are attacked, the women's epithelial tissue is the problem instead. But Sarah's fertility is proof of how she can beat it. Paul figures out that Coady is trying to isolate and weaponise it. But someone else seems to be calling the shots.

Rudy sneaks back into camp while his driver is placed under arrest. He lets out one of his brothers, Miller, and hands him a weapon so they can take back the base. Paul is being advised by Sarah that they need to get out of there and Paul assures her that help is on the way. But the conversation is interrupted by Mark, who says that patrol came back seemingly without Rudy and that upon checking the SAT phone, he has found that they got a call from Arlington. Paul then calls his contact and as he lies, he knows that he has been betrayed and no one is going for them. Paul and Sarah must escape now, and Mark tells them to make this right. Paul gets into a fight with Miller and has to snap his neck but is badly wounded by Miller's knife before he makes his kill. Paul leads Sarah to a tunnel and locks it so she can't get him to leave with her. He tells her to follow the tunnel to the garage, grab jeep, and leave. He tells her that he loves her and leaves. Sarah, having no other choice, heads off.

Paul is shown to be in Coady's office with the science. Coady and Rudy walk in and when Paul is shown to be refusing to leave his course, Coady shoots him repeatedly. Paul is then revealed to have a grenade, which he activates. Rudy and Coady run out, possibly/probably with enough time to get to safety, and Helena picks up a disorientated Sarah following the explosion.

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Helena may be back in her cell following her mercy-killing of Castor's lab rat but what she doesn't know just yet is that Sarah is now in the cell right next to hers. Meanwhile, Cody and her science team will get to work on the baby remains to try to isolate the genome so they can resolve the defect in the Castor clones.

Sarah tries to figure out a way out but her yelling alerts Helena to her presence but Helena is holding a grudge, believing that Sarah is to blame for the captivity she has faced for all this time. Sarah tells her it was Mrs. S's doing and that the decision had been made from a place of trying to protect Kira, though Sarah admits it was a bad call.

Gracie arrives at Art's apartment to turn herself in and ask for help, as Sarah had given her Art's card to go to him for help if ever she needed anything. Gracie shares some of what she has been through regarding her mother but she doesn't want to talk about Mark. She admits she lost the baby and this is the reason she was cast out.

Cosima calls Felix for a pep talk before her first found-via-Internet date. Her date comes in the form of a hot holistic dealer named Shay. Despite Cosima's nerves, the two hit it off quite well. Cosima opens up a bit about Delphine and by the end of the night, Cosima's getting a massage and smooches.

Sarah gets taken out for questioning and on her way out she calls out to Helena to tell her that the Castor clones are their brothers. Helena also discovers that the bars on her cell door's window are a bit loose and it turns out she's been stashing butter from her meals behind a loose stone in her cell wall. Dr. Cody forcibly takes some samples from Sarah, declaring that her biology is too special to ignore. Cody is also a complete creep, as she tells Mark that his marriage wasn't real and he gets his first "log book" in which all intimate contacts are to go into.

Gracie is brought by Mark to Mrs. S and Felix so they can look after her as she is not about to essentially be deprogrammed. She just wants to experience all the norms of being an 18-year-old girl. Art and Mrs. S are also starting to get concerned about Sarah's disappearance.

Sarah's trying to pump Helena for information, knowing that she's bound to have counted the number of people on the base and have scoped out the layout. But their conversation is interrupted by the arrival of Paul, who tells Sarah all she had to do was stay away. He is so obviously still carrying a torch for her. Paul claims that all he is doing for the Castor clones is the same thing Sarah does for the Dyad clones. Sarah pleads with him to let them go but he says it's out of his hands. He claims that all they're doing is to try to save the Castor clones and that Helena is no saint either. Helena seems to now believe what Sarah says, as Paul admitted during the conversation that he was the one that Mrs. S sold Helena out to, saying it was either her or Sarah.

Sarah's still trying to mend things with Helena so she opens up about being an irresponsible mother when Kira was first born. She had a drug problem and one day, she finally left town with Vic to prove Mrs. S wrong because one day Mrs. S refused to let her see Kira. Sarah is wracked with guilt and now is determined to make for it to Kira and has a mama-bear complex about saving all the clones. Now Helena is finally willing to let Sarah in on her plan!

We learn a bit more about Mrs. S, whose past love had a drinking problem and he died because of a drunken row. But the conversation with Felix is interrupted as Gracie arrives in some of Sarah's punk-rocker clothes and declaring she wants to go out. Felix is able to reel Gracie in so they can do the drinking in the safety of their own home. The first order of events is to give her a cocktail, introduce her to some rock music and teach her to dance. But the fun is short-lived, as Gracie collapses to the floor, feeling sick and suffering from abdominal pains.

Sarah tries to fight some of the male guards when they come in due to her refusal to hand over her lunch tray. She takes a few hits and ends up in the medical centre where she swipes a tool off the tray when the physician's back is turned. Turns out it was a pair of tweezers, which Sarah must swing over to Helena through the front of her bars, tied to the end of her belt, but time it so as not to be seen by the moving surveillance cameras. Next, Helena picks the lock on her cuffs and breaks out the bar. She's also lubed herself up with her stash of butter so she can squeeze through the bars. She then messes up the camera so no one can look in. When the guard comes in to check on it, Helena impales his head through a spike on the wall and spits on his corpse for good measure. In the most shocking turn of events yet, Helena leaves Sarah behind and declares that they are now even.

Paul is looking through the base and appears to be having doubts, as he examines Parsons' body and the place where he had been strapped up beforehand. He also finds Parsons' log books with the hair samples and detailed descriptions of all the different womyn he had hooked up with. While Helena continues to scale the walls of the base to make her escape, it's then that we learn that whatever defect the Castor clones are suffering from can be spread as a sexually transmitted disease. The Rudy/Seth hook-up girl is sick and now so is Gracie. Helena's having guilt over leaving Sarah behind but since the base has been put on high alert, we won't know just yet if Helena will escape into the night or come storming back in with some grand, elaborate rescue plan. Perhaps both!

Orphan Black airs on Saturdays on BBC America at 9PM.

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Surprise! Mark's not dead! Despite his protests, Sarah insists on helping the wounded Mark out of the cornfield just in time to get him away from Gracie's mother's Prolethean goons. Mummy Dearest is bringing Gracie back to the compound and Gracie must tuck away her wedding ring for safe keeping.

As for Helena, our favourite bottle-blonde trickster ties off her arm to slow down the sedative being inserted into her system so she can go wandering around the base. She makes it back to her room in time with a the bone from a steak which she proceeds to gnaw on to mould it into something special.

Alison gets paid a visit by a henchman of the local drug kingpin because Ramon, whom they bought the drugs off of, didn't actually leave to go to college. He made a buy-and-dash and is now missing.

Sarah has helped Mark into an abandoned house where she's able to dig the bullet out per his instruction and sews him up. It seems Mark had hoped to trade what he found for his freedom but he tells Sarah that what he found instead was just a bunch of junk. Sarah goes through Mark's stuff and swipes his motel key and gun before calling an ambulance for him, as he has just passed out.

We see little from Cosima save for her wearing a Yeti-like sweater that Felix hates. Felix advises her not to do so much pining after Delphine and to move on with her life. Yeah, Cosima! Get some honest loving this time around! Felix thinks Cosima getting on a dating app is in order to find her next scratching post but Cosima is reluctant to do so.

Alison gets to meet with local drug kingpin, and it's Jason Kellerman, her ex-boyfriend from high school! Thankfully, they're on friendly enough terms and Jason doesn't seem too upset that she has moved in on Ramon's old action. We also learn that Alison's mother always like Jason. That's certainly interesting. He wants to know why she's doing this but she doesn't feel like sharing. He respects her affinity for business and admires her savviness regarding the soap idea, which he calls "genius." She promises to double his profit on the stock, if he'll just give it back first.

Sarah makes it back to Mark's motel room and goes through the journals and other papers that were recovered by Gracie. She relays the news that Mark was undercover with the Proletheans and what's news to us is that the Prolethean patriarch Henrik Johannson was Professor Duncan's lab assistant during the earliest days of the experimentation. It also looks like Gracie's mother was responsible for carrying the clone embryos in the earliest days.

Gracie ends up miscarrying and her mother is enraged by this. She spouts some religious dogma and kicks Gracie out of the compound.

Mark corners Sarah in his motel room and she shares what she has figured out after sharing the information with Cosima who could decode the scientific chicken scratch in the notebooks. The reason why there were no samples is because Johansson and his wife used the original tissue samples to make a baby. All they need to do is find him, get some tissue samples and they'll be good to go. Mark knows that Johnasson had mentioned a son so he tells Sarah where to drive. The son died so now Sarah must dig up the grave of a dead baby.

Remember that bone? Well Helena has chomped it down into a pick for the lock on her cell door. How resourceful! Once she's broken out, she discovers a castor clone named Parsons who looks to be in bad shape. He's restrained and Helena discovers that the top half of his skull has been removed and his brain exposed with some electrical nodes sticking into it. He begs her to kill him so he can be put out of his misery. Her scorpion friend advises her to not risk her one chance at escape on someone that is already dead. Helena gives a moving speech to Parsons about them both being abandoned by their families and left to suffer. It's sad to see that Helena has bought into the BS she's been fed about her sestras not caring about her. In perhaps the "sweetest" death we've seen yet, Helena takes a scalpel to Parsons' brain. Cody comes in calling Helena names but Helena snaps back by calling her a liar who claims to love her boys but really she's just a "shit mother." Bravo, Helena, bravo!

But there's one last twist to be had, as Rudy shows up to check Mark for a pulse just as Sarah has gotten the baby's remains up and out of the ground. She tells Rudy what's in the box and when he goes to open it, she whacks him with a shovel and takes off running. She may not have the box but at least she's alive for now. Unfortunately, Rudy's still awake so he comes stalking after her in the barn. Just as Rudy's preparing to kill Sarah, Mark comes in declaring that he is the superior officer and order Rudy to stand down. Rudy knocks Sarah down and keeps his stand-off going with Mark who dares him to kill the both of them if he thinks he's such a tough "little brother." Rudy stands down but Castor clones don't leave loose ends, so it looks like Sarah's going to have take a little trip...

Orphan Black airs on Saturdays on BBC America at 9PM.

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New York, NY – May 7, 2014 – BBC AMERICA announced today the renewal of its award-winning original series “Orphan Black” for a fourth season.  The drama starring Tatiana Maslany has been re-commissioned for a 10 episode order from Temple Street Productions and is slated to air in 2016. Hailed as “utterly original and spellbinding” by The Daily Beast and “a multi-layer cake of performance” by The Los Angeles Times, season three is currently airing and will finale on Saturday, June 20 at 9PM ET.

Sarah Barnett, BBC AMERICA President and General Manager, stated, “’Orphan Black’ is an entirely fresh and original show that stands out even in today’s crowded TV landscape. Tatiana Maslany’s buzzy, virtuoso performance headlines a brilliant cast and we are delighted to let our vocal and passionate fan base know that this wildly entertaining ride will continue next year.”

Co-creators John Fawcett and Graeme Manson said: “We’re thrilled to be moving forward with season four of the Orphan Black saga. Huge thanks to our partners Temple Street Productions, and to BBC AMERICA and Space. We’re extremely excited about where our story lines are going, and look forward to riding the rollercoaster with our amazing fans #CloneClub.”

The series has received considerable critical and award attention. It earned a Peabody Award in 2014 and lead actress Tatiana Maslany has received two Critics’ Choice Awards for Best Actress, a TCA Award for Individual Achievement, a Young Hollywood Award, a Gracie Award, two Canadian Screen Awards and Golden Globe®, SAG and People’s Choice nominations.

The cast includes Tatiana Maslany, Ari Millen, Jordan Gavaris, Dylan Bruce, Maria Doyle Kennedy, Skyler Wexler, Kevin Hanchard, Evelyne Brochu, Kristian Bruun, Ksenia Solo, Michiel Huisman, Josh Vokey, Zoé De Grand Maison, Justin Chatwin and Kyra Harper.

“The growth of ‘Orphan Black’ and the amazing response from audiences over the past three seasons has exceeded our expectations and we’re delighted to be able to continue the journey in season four,” commented David Fortier and Ivan Schneeberg, executive producers of “Orphan Black” and Co-Presidents of Temple Street Productions. “We are incredibly proud of our talented cast and crew and grateful for the support from our broadcast partners at BBC AMERICA and Space.”

“Orphan Black” is produced by Temple Street Productions in association with BBC AMERICA and Space. Executive producers are Ivan Schneeberg and David Fortier of Temple Street Productions, Graeme Manson and John Fawcett. “Orphan Black” is distributed by BBC Worldwide.

“Orphan Black” Season Three Audience and Social Media Highlights:

• The AMC Networks roadblock of the “Orphan Black” Season 3 premiere garnered the largest ever audience for the show, with 1.9 million viewers. 
  • BBC AMERICA accounted for 60% of the audience with 1.1 million viewers on premiere night.
  • Source:  Nielsen, Live+3, 4/18/15, Orphan Black Ep 301 (BBCA, AMC, IFC, SUND, WEtv), P2+ gross impressions.
    Includes premiere night encore on BBCA.

• With a Median Income of $77K, “Orphan Black” ranks as the 2nd most affluent drama on ad-supported television in the 2014/15 season, behind only AMC’s “Mad Men.”
  • Together, AMC and BBC AMERICA account for the Top 4 most affluent dramas on ad-supported TV this season: “Mad Men,” “Orphan Black,” “Better Call Saul,” and “Doctor Who.”
  • Source:  Nielsen, Live+SD, 2014/15 STD (9/22/14-5/3/15), Broadcast & Ad-Supported Cable: Primetime Original Dramas (excludes repeats), Median Income (P2+).
• “Orphan Black” continues to show tremendous social media engagement, ranking as the #1 basic cable drama on Twitter and Tumblr in its premiere week, and continuing to rank among the Top 10 cable dramas on both platforms each week thereafter.
  • Twitter Source: Nielsen SocialGuide, week of 4/13/15, basic cable program rank based on unique authors.
  • Tumblr Source: ListenFirst Media, week of 4/13/15, basic cable program rank based on engagement (notes & tags).

About BBC AMERICA

BBC AMERICA delivers U.S. audiences high-quality, innovative and intelligent programming. Established in 1998, it has been the launch pad for talent embraced by American mainstream pop culture, including Ricky Gervais, Gordon Ramsay, Graham Norton, and successful programming formats including ground-breaking non-scripted television like Top Gear and top-rated science-fiction like Doctor Who. A joint venture between BBC Worldwide, the commercial arm of the BBC, and AMC Networks, BBC AMERICA has attracted both critical acclaim and major awards including four Emmy® Awards, five Golden Globes® and 12 Peabody Awards. The channel attracts one of cable’s most affluent and educated audiences and is available on cable and satellite TV in nearly 80 million homes. It broadcasts in both standard and high-definition, with content available On Demand across all major digital platforms. Online, www.BBCAmerica.com is the place to go to dig deeper into pop culture with a British twist. Follow us on www.twitter.com/BBCAMERICA.

About AMC Networks Inc.

Dedicated to producing quality programming and movie content for more than 30 years, AMC Networks Inc. (NASDAQ: AMCX) owns and operates several of the most popular and award-winning brands in cable television. AMC, IFC, SundanceTV, WE tv, and IFC Films produce and deliver distinctive, compelling and culturally relevant content that engages audiences across multiple platforms. The company also operates BBC AMERICA through a joint venture with BBC Worldwide.  In addition, the company operates AMC Networks International, its global division. More information is available at www.amcnetworks.com.

About Temple Street Productions

Temple Street Productions is a leading producer of television, film and digital media content based in Toronto and Los Angeles. A selection of Temple Street’s past and present projects include, Orphan Black (BBC AMERICA, Space), X Company (CBC), Being Erica (CBC, SOAPnet), The Next Step (Family Channel), Wingin’ It (Family Channel), Say Yes to the Dress Canada (W), Million Dollar Critic (W, BBC America), Cook’d (YTV), Over the Rainbow (CBC), Recipe to Riches (CBC, Food Network, Global), Cover Me Canada (CBC), Billable Hours (Global, Showcase), Canada’s Next Top Model (CTV), How Do You Solve A Problem Like Maria? (CBC), Queer as Folk (Showtime, Showcase), and Darcy’s Wild Life (NBC, Discovery Kids, Family Channel). For more company information www.templestreetproductions.com twitter @TempleStreet or connect with us on Facebook.

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Sarah and Felix have quite the task on their hands: disposing of Seth's body. Of course the timing couldn't be worse for Art to show up at Felix's flat. He knows they're up to something so he comes barging in and finds the body and Sarah's first response is to truthfully tell him that they weren't the ones to kill him. So now it's time to Skype-call Cosima (and Scott). They determine that the male clones must have a neurological defect which is why they're after the original genome. Cosima then declares that they need to examine his brain... because science! Art also lets Sarah know that they've been able to track down a lead regarding Mark.

Mark's got his arm bandaged up after burning off his tattoo and despite a delay in consummating his marriage, Gracie's encouraging words coax him into finally doing the deed. It's also time for him to start opening up. He shares about his time in the military academy and how despite all the other boys and their chasing tail, he wanted to wait. He says that her father had stolen valuable scientific material and he was assigned to recover it. Mark says they're free from the Proletheans but now he needs to get free from the military. He has to recover his material, hence their hiding out in a seedy motel.

Art and Sarah catch up to the midwife that helped artificially inseminate Helena. They learn that Helena is impregnated by that creepy Prolethean leader and that a fertilised egg has been implanted into Gracie, too. Turns out Gracie is strong at bluffing which helps her to recover the stolen science material that was being kept by Willard Finch, a contact of the Proletheans.

We've got an update on Alison, too. She's working her sales skills, with the help of Donnie, to simultaneously campaign and let their clientele know who the new pill-pushers in town are. She's selling the pills by hiding them in handmade soaps. Her competitor tries to bribe Alison off the warpath by offering her a good deal on a house in the nicer district but Allison will not be so easily swayed.

It also seems as if Art was in love with Beth and that's a huge reason why he's pushing so hard to help the clones. He feels guilty for seemingly abandoning Beth in her hour of need.

Paul brings Rudy back to the base and Dr. Cody is indeed the "mother" to the clones (at least by name, of course). She's furious about Rudy putting Seth down though Paul insists that this was entirely necessary. Dr. Cody is concerned about the glitches in their weapons program though it's unclear yet just what it is she's referring to. Paul thinks it's best that Rudy stay on the base but Cody insists that she needs him back in the field.

Cosima and Scott head to Felix's place where Cosima takes a saw to Seth's skull to get a look at his brain. Oh and it's worth noting that she learned how to do all of this from some Youtube videos. That's our favourite mad scientist!

Rachel is on the mend, though she's disappointed to learn that Delphine is now the one in charge at Dyad instead of her. She's being given cards of symbols to identify and unfortunately for the tester, she doesn't recognise the two-headed horse as the symbol of Castor. It turns out Rachel knew a lot more than we originally thought.

Art and Sarah track down Willard Finch in search of Mark and Gracie. With a bit of arm-twisting, they get him to admit seeing Gracie. Mark is discouraged when Gracie returns with the stored information that doesn't actually include any biological samples.

Rudy pays a visit to Helena, who dubs him "the ugliest Mark" of the bunch. They exchange some fighting words before Paul comes in to pull Rudy away to be debriefed before he'll head back out in the field. Paul tries to apologise to Helena, though Helena promises that one day she'll kill them all, and even throws in some bullet sound effects to top things off.

As luck would have it, Sarah stumbles across Gracie in a diner. She makes it clear that Gracie needs to do what's best for her/Helena's baby and tell them where Mark is. Mark is torturing Willard Finch to get more information but while he's away, Gracie's borderline-psychotic and entirely puritanical mother shows up take her back home. She got Gracie's location from the banished midwife.

Sarah is at Willard's farm to find Mark but while searching, Cosima calls from the lab to let Sarah know that she and Scott have made an incredible discovery. The Leda and Castor clones are siblings, as in, the original donors from which the clones were spawned were sister and brother. But this can't be discussed further just yet because Sarah's got a show-down with Mark. He's furious to learn that Sarah told Gracie about Mark being clones (so in her mind, an abomination). Sarah tells him about their being siblings but Mark doesn't care. He wants to run off and mend fences with Gracie. But when he runs outside, Sarah must watch in horror as Gracie's mother shows up and offs Mark with a big gun.

Orphan Black airs on Saturdays on BBC America at 9PM.

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Rudy's been broken out of his holding cell by Seth and their first order of events is to try to "share"a girl who gets a whole lot more than she bargained for.

Sarah and Kira are reunited with Cal, who has one fabulous new home that's both spacious and cozy. Sarah and Kira both seem to like it, but Sarah doesn't seem too quick to trust that she'll be able to even consider a "normal" and domestic life any time soon.

Surprise! The Castor clones and Paul know each other! They seem to be under his authority and he has to run some tests on them, though it's not entirely clear why. Probably all part of protocol, I suppose. Rudy aces the test, but Seth seems to struggle on one of the questions. Rudy blames it on the alcohol consumption and sleep deprivation. They have their orders, clean up and clear out. They're to be extracted and brought back to base. Doubtful that Rudy will go gently into that night, though. He hardly seems the type to take orders.

Alison still wants to run for office but with Donnie's recent unemployment, she'll need to find another way to raise the funds to afford her campaign. When she learns that the local prescription-pill drug dealer is going out of business since he'll be headed off to college, our favourite little housewife gets an idea about how to make some money.

Sarah gets a call about Rudy's escape. She's still determined to find Helena and wants to make sure that her family is safe. She and Cal are rekindling things but let's be real here, Sarah's relationship with Cal is the least interesting relationship she has on the entire show. As for poor Helena, she's still in captivity and being waterboarded by Miller and whomever else is running the show. But the waterboarding must come to a halt because the bloodworm came back and our favourite little mad hatter is pregnant! Dr. Virginia Cody is the one that halted the waterboarding/taking of blood and saliva samples and seems to be the one in charge. But who exactly is this mysterious Doctor? Is she the "Mother" of whom Paul referenced while speaking with Rudy and Seth?

Mrs. S is still on the mend after her run-in with Seth and Felix checks in on her well-being. He also assures her that Sarah will come around eventually, referring to the revelation that Mrs. S was involved in Helena's capture.

Cosima and Scott meet with Dr. Nealon, the new scientist in charge at Dyad under Cosima's regime. Nealon tells them how everyone is after the original genome and that the Duncans found two lines of clones (Leda/Castor) but never disclosed the original donors. The Duncans were the ones to start a female and male lines. Only the female line of clones was brought to Dyad when the military shut the project down (could misogyny be the reason that the military only kept the male clone projects operational?)

Art lets Sarah know about a call (from the woman who met Rudy and Seth). Thankfully, she's still alive and seemingly uninjured albeit freaked out. Turns out Rudy wasn't happy about this woman rejecting Seth's advances. She got a lock of her hair ripped out (a trophy of sorts) and all her personal information off her ID was written into their little notebook. Weird weird weird. Art warns Sarah not to impersonate an officer again but we can't help but love our mama lioness' gung-ho ways. She will do whatever it takes to protect her family.

Alison and Donnie meet with Ramon since Alison wants to buy out his business. She's one hell of a convincing saleswoman; how was she not in this line of work already? Ramon agrees to sell but Donnie's getting skittish. Allison insists that this is about their kids but also notes that her newly acquired clientele will also be voters. Donnie is in awe of her scheming ways.

Rudy calls Seth up to tell him that there's no way he's going back to base. He knows Seth is glitching and Seth admits it's been happening for a week and it's only getting worse. When Rudy makes the call, he is shown to be breaking into Felix's apartment and he says they have no choice but to "finish the job."

Cosima advises Sarah to get back in touch with Mrs. S so she can get in touch with Paul as it's their best lead on getting to Helena. The centre is trying to run the same exercises on Helena as what was being done earlier on Rudy and Seth but she's distracted by her scorpion friend and wants to know where the mangoes are. Oh, Helena. Her sass is just as strong as ever. Dr. Cody is trying to manipulate Helena by claiming that her sestras sold her out. Helena doesn't believe her though. Cody says that the Leda clones are more resilient to both nature and nurture than the Castor clones. Surprise, surprise, the weaker sex is defective.

Sarah goes to ask Mrs. S where Paul is but apparently Paul has gone dark. Sarah isn't about to dish out forgiveness anytime soon. Paul makes an appearance and much to our surprise, it's to visit Cal. He knows some dirt on Cal, who is actually a bit of a war profiteer. He only got out of his misdeeds after he got rich. These two had a deal that would buy Sarah space and time but unfortunately, Sarah has been refusing to take either. Paul tells Cal that Sarah needs to stop digging into Castor for her own safety.

Sarah and Kira have gone back to Felix's place and the running water of the tub means that they don't hear Cal's call. Poor Kira crawls into her blanket fort and lo and behold, Rudy's there and sporting some new red X's on his eyes. Is this a new Castor fashion statement? He holds Kira at gunpoint and tells Sarah he wants Duncan's research and the tissue samples of the Castor original. Say what??? Cal shows up downstairs to try to help Sarah and Kira but Seth is playing watchdog. But then he starts glitching. Sarah was already to piece together that Seth was sick (since they've got Cosima, their own sick clone). Rudy runs off when he hears Seth's hollering. In a surprising turn of events, he shoots Seth in the chest to put him out of his misery, bidding him goodbye with an "I love you." We really thought that Cal was the one who'd get shot but that's what we love about Orphan Black; we're kept on our toes!

Sarah ends up bidding goodbye to Kira, deciding the best course of action is to send her off with Cal to Iceland. Why didn't she do this sooner? Mum needs to kick butt and destroy their enemies like the badass superhero she is before she can do anything remotely domestic. "Now we find Helena and finish this shit." How will Sarah do this? She sets her sights on finding Mark. The Prolethean clone is currently holed up in a motel with his new wife, Gracie, but first he needs to burn off the tattoo from his arm. This tattoo is one that all the Castor clones have. Spoiler alert, it's a tracking device!

Orphan Black airs on Saturdays on BBC America at 9PM.

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Helena has a dream of a BBQ baby shower in which all her sestras (plus Kira and Felix) are present. But it quickly turns to a nightmare in which a scorpion crawls out from her dress up towards her face until she wakes up trapped in the box that is her new home (till she can make her escape, of course). There is an actual scorpion in there that talks to her, and of course she talks back, and it's quite the pep talk to encourage her.

Delphine rolls up her in her fancy new car to meet Sarah, Felix, and Kira. She brings Sarah in to tell her about Rudy, the big-bad Castor Clone with the mohawk and the facial scar. He was caught in cahoots with another Castor clone (that got away) and together, they tried to kidnap a new clone that we don't see apart from some steamy surveillance footage of her hooking up with Rudy in an elevator. Now he'll only speak to Sarah.

This facedown is rather intense. He says Sarah's a legend and makes some not-so-thinly veiled threats, namely that she should count her sisters. A quick few phone calls makes her realise that everyone is fine for now but Helena is missing. Delphine has even more warnings, saying that Top Side will be sending people in to check Dyad and she needs Sarah to play Rachel (who is still in recovery and getting the pencil removed from her eye).

Alison decides to run for school trustee because she doesn't care for the current candidate's plans to redraw district lines, which would mean Alison's children would have to change schools. Oh and Donnie got fired because he finally told his boss off, and he's very excited about it.

Mrs. S comes home only to get attacked by one of the Castor clones. Of course she's no wilting flower, so she gets a few good shots in as well as a stab to the leg. This clone's named Seth and he's got a pornstache. He wants Duncan's research but she's not singing and surprisingly, he leaves her alive. Sarah and Felix find a bloodied Mrs. S and she tells them the truth about handing Helena over, as part of Paul's demand, and Sarah is livid. Mrs. S did what she thought was best for the good of the clones but it'll take more than that to win Sarah's good favour back.

Delphine's getting shadier every minute as she meets with one Dr. Nealon and tells him to put the good of Project Leda as a whole over the originals. Delphine seems like she's not so keen to play favourites anymore. Delphine then pays a visit to tell Cosima about her promotion. She says she'll keep her promise to love all of Cosima's sisters equally, but in order to do that the relationship is going kaput. Cue the box of tissues!

Alison and Donnie need to sort out their finances. They have some struggles to face in that area but now Donnie is adorably supportive of Alison's desires to run for board trustee. We are entirely here for this dream-team duo. Also speaking of duos, Cosima is teaming up with Scott to try to crack the code in Duncan's book. She hasn't even told her sisters about her plans.

Sarah decides to help Delphine fool the Topsider cleaner Ferdinand so she can leverage Delphine's help in getting Helena back. Of course Felix is there to aide in this charade. Ferdinand is played by James Frain so there's an automatic creepiness (that's Frain's specialty). The plan was to have Sarah-playing-Rachel say hello and leave but he wants to speak to Delphine and Rachel together. Once "Rachel" drops the bomb that Helena's been kidnapped, Ferdinand wants to question Sarah, whom he has led to believe is in captivity. But the "Sarah" Ferdinand meets is actually Alison in a strange wig. Ferdinand feels up "Sarah" to "examine" her and "Rachel" manages to deflect it by slapping her for getting mouthy. Sarah was supposed to have her ovaries removed (they don't want the clones breeding). But Ferdinand pulls "Rachel" aside and says he wants to discuss Helsinki, while "Rachel" just wants to discuss Helena, saying that her being out of their hands is also a risk.

Rudy's doing naked pull-ups (the guards have dubbed him "Naked Buddha") and we're not mad about it. We're betting this routine behaviour is part of his military-rooted upbringing.

Delphine pays a visit to the recovering Rachel, who is having difficulty getting her words out. Delphine's grim new demeanour rears its head when she presses on Rachel's wounded eye to make her talk. Delphine wants answers regarding Ferdinand and Helsinki.

Ferdinand has showed up to "Rachel's" and finally the truth comes out. Rachel and Ferdinand had offed six clones in 24 hours in 2006. He's already putting action in place, as armed men are heading out to take out Alison, shortly after which more of the clones will be taken out. Rachel had done this with all because she wanted to be a mother. She was supposed to get Sarah's ovary(s) and make off with Kira. Sarah tries to call Alison and warn her but Ferdinand comes in and interrupts her. He's still getting suspicious but Sarah's able to fend off his curiosity by whipping off his belt and whipping him with it. She then gets on his back as he lies on the bed and begins strangling him with it.

The strangulation is interrupted by Delphine, who has learned all about Helsinki. Ferdinand's higher-ups wouldn't be too pleased if they learned an employee had conspired with one clone to kill the others. Delphine now has leverage on him and makes him text Alison's would-be killer to call the plan off.

As for poor Helena, she's still talking to her new friend, the scorpion. She's finally let out by Miller, the military (well, most obviously military judging by that outfit) clone. By Helena's not the only one getting out of boxes. Seth AKA 'pornstache clone' shows up where Rudy is being kept and kills a guard before letting him free. They also share a hug. Aw, how broclonemantic!

Orphan Black airs on Saturdays on BBC America at 9PM.

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One of the highly anticipated plot developments for season 3 of Orphan Black is that of the revelation of the male clones of Project Castor. Actor Ari Millen was introduced in season 2 as Mark but in the season finale, it was revealed that Mark was but one of many! Ari joined us for a conference call interview and teased about his core group of four clones and discussed the challenges of taking on this array of characters.

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Being that the Castor clones were raised self-aware, how does that change who they are to each other and their brotherhood when compared to the Leda clones?

Ari Millen: The fact they're self-aware, meaning that they grew up together, they have a very close-knit, wolf-pack of a military upbringing. Just like any set of brothers, there's a real closeness, they like taking the piss out of each other, they can be vicious, but there's also a lot of love there compared to Project Leda who are just learning who they are. As the season progresses, Project Castor is definitely going to hit the ground running as far as making it very difficult for Project Leda because they have something that 'we' want and because we know who we are, we have a certain better understanding and are almost more cohesive, whereas Project Leda is not only figuring out who they are but who they can and can't trust. The Castor clones are a very strong foe right off the bat.

Who is Paul to the Castor clones? What can you tease about what we'll see of him and the male clones and do they trust him?

Ari Millen: Oh man. Without giving away spoilers, I think at the end of the season, we now know that Paul knows more than he was letting on and I think that he might have been playing both sides. As the season progresses, we'll find out more of what Paul knows and more of his motives and who he is to Project Castor.

With the Leda clones, there's a spectrum of sexuality. There's Cosima, who's a lesbian, and we have Tony, the trans clone. What would you say is the spectrum of sexuality for the Castor clones?

Ari Millen: They're ultra-hetero. [Laughs]. Oh man, this is a good question. I think part of how they wanted to position Castor this year was as this sort of testosterone-menace and I think that aspect plays itself out through their ultra-sexuality. It's a sort of frat-boy mentality, but that is just one part of them. But on the complete opposite side of that, you have Mark who is almost virginal and almost innocent. There's some interesting dichotomies with them but the sexuality will play a big part for this season, that's for sure.

Since you were a part of season 2, you've had a chance to see Tatiana in action. What was the biggest challenge for you since you've been having to do so much more than you did last season?

Ari Millen: I think the biggest challenge was, since Castor is self-aware and they grew up together, finding the differences within the similarities. Finding the little nuanced things that make Rudy Rudy, that make Mark Mark, that make Seth Seth. On a multiple Clone day when I was playing more than one of those Castor guys, just making that switch. Because they're so similar, trying to drop one and pick up the other, depending on how much sleep I got the night before might've been more challenging. It was a really fun process of trying to find these guys within their brotherhood.

Which of the Castor Clones would you say is the most dangerous that you wouldn't want to encounter in a dark alley?

Ari Millen: I think the obvious answer is probably Rudy but, I think Rudy to a certain extent wears his heart on his sleeve. I think Mark is the more dangerous one. He's very good at hiding what he's doing and you don't want to cross him. You don't want to threaten what he's going for because he will make you pay.

What is going on with Mark and Gracie? Will we see more of them?

Ari Millen: Yes, Mark and Gracie have just eloped at the end of this last season and they're trying to make their escape. They're trying to hide from the Proletheans and now with this new revelation of Project Castor, we find out that they're also looking for Mark as well.

What kind of advice did you get from Tatiana about playing multiple characters?

Ari Millen: The first thing that happened was, she knew probably a day or two before I did that it was gonna be me, so the first thing that happened after I found out, and it was still a secret to the rest of the cast and crew because we hadn't gotten the season finale script yet, she just sort of came up to me and gave me a little rub on the back and winked at me and said, "Congratulations." That's how it's been for us this season too. Acting is such a private and personal process that it's never something that you impose on anybody else. For the most part, she's present and there when you're in the scene together, but if I ever had a question about, "How do you act with those tennis balls so well?" she would be there to answer it. But I find for me that the best lessons that I can learn are just from watching people who are good at what they do, do it. So that's what I would do. When I was on set and she was on set, I would just watch her do her stuff and see the little things that she was doing and say, "Oh hey, that's brilliant. What a great idea." Then I would go on my turn and try that little trick out and if it worked for me, I would keep it and if it didn't, I wouldn't. For the most part, it's a visual process. She's very supportive.

So did you find any tricks that you used to get into the headspace of each of the clones? She has certain playlists for hers so did you have anything that you did to inhabit those guys?

Ari Millen: At the end of the last season, it was more of a reveal but no one knew who they were at that point other than John and Graeme who knew what they wanted. We got these wicked visuals, like of Rudy with the scar and the mohawk, and Willard the military guy. That's what I had to sit with during our time off so when it came to creating the guys, it was a visual 'in' for me. I would put on my wardrobe for the day, I would go to hair and makeup and they would put that on me for that day and I would look in the mirror and drop into the decisions that I had made. "Okay this guy is like this and this is what he does." It was a very visual process for me and if ever I found I was coming out of it, I would go find a mirror and settle, take a breath, and just look and drop back in. It was a very different process for me than normal but it still worked. It's interesting, I had an acting teacher once say that whatever works is good. There is no right answer. If it works, it's good.

There's some major differences between season 2 and season 3 for you. In season 2, you're acting with other people and you've got people to play off of. In season 3, you're getting to do scenes with potentially just yourself. What's been the most difficult or perhaps most fun element of filming things like that?

Ari Millen: Well I'll start with the fun. Being on a show like Orphan Black, you get put in a lot of really crazy and exciting scenarios. The fact that my screen time got multiplied this season only meant that I was gonna get to do more fun and exciting stuff. With that came the challenge of acting with myself. The challenge was learning how to plan out the scene for both characters at once. You shoot the first half of the scene as one character and then you switch over and do the scene again as the other character. The trouble could be, if not handled properly, if I only pay attention to the first character I was playing and forgot about the other guy then the decisions I made, I would've been stuck with when we flipped the camera. The big challenge this year, especially working with my clone double Nick Abraham and the director, we would just talk it out and plan it and make sure that if it was a scene with Rudy and Seth and I'm playing Rudy that we make sure that Seth does 'this' at this point because this is the moment that I want to create for Seth. There was a technical side of trade planning that wasn't organic to me so it was a very interesting learning curve to do this season.

Were you able to put any input into the characters? Did you put any of your own spin or nuance into their personalities or looks?

Ari Millen: What's great about this show is that there is a real fluidity to the script writing. There's a lot of frustration with only learning the next episode part-way through the episode that they're shooting but that also lends itself to being organic and switching on the fly. Certainly Graeme and John and the rest of the writers always were happy and interested and inviting for our input. What I found really cool too was a lot of the time, I would come to them with this instinct that I had about it and then they said, "Don't worry, we already got you covered in the next episode," so it was like we were on the same wavelength with a lot of stuff. I think one of the greatest things about this set is that it's a collaboration. There's not one dictator at the top, it's a collaborative process.

In the season 2 finale, that moment where Mark and Gracie are getting married is huge. Once you knew what Mark really was, did that make you modulate how you played that scene? Was there more menace to it or was it more protective? How did you want us as an audience to look at Mark in that moment?

Ari Millen: One of the things I found most exciting and I enjoyed the most about this season was the rediscovery of Mark because I didn't know what we all know now, last season. Specifically in that scene, I was still in the mindset of who Mark was from the point of view of last season. I think often it's dangerous to play the end in that. I think it was more important just to see him getting what he had always dreamed of. That was the most important thing. That's what he always wanted. He always wanted that sort of fairy tale, of disappearing out of wherever he came from and going forward with the white picket fence, the wife, and hopefully soon, children. So in that moment, it's just pure happiness for him. I don't think he was thinking about anything else other than, "This is the woman I want to marry." Castor doesn't even factor in at that point.

For that really wonderful reveal at the end of the season, what was the conversation with Graeme and John on which visage to show to Sarah in that really intense moment?

Ari Millen: The little that they knew about Rudy at that point was that they knew they wanted him to be a little bit unhinged. I think that was really concrete direction and then we just played. We probably did ten different takes in ten different ways of that reveal and that's the one they chose. They were all similar but different.

Will we see conflict within the Castor clones?

Ari Millen: Yeah, they're brothers, they fight. That's kind of a generalised way to say it. Certainly Mark has his own ideas of who he wants to be and where he sees himself going. His brothers in Project Castor are gunning for him. They're not going to let him go without a fight. There's gonna be some conflict there, there's gonna be some butting of heads, and fights. Within the other guys, they're reaching a certain point in their life, and I'm gonna try to be as vague as possible so I'm not giving spoilers, but they're reaching a certain point where something is becoming very very important so there is a certain amount of stress that they're all under. The bickering will start but for the most part, they're a cohesive group and they're all working towards a common goal and even though there's fights, there's ultimately love and family and that's the most important thing.

Which clone did you find the most intense to play?

Ari Millen: I guess the answer is Rudy. What's really fun about Rudy is that it's often a game to him to try to push people's buttons and knock people out of their comfort zone. We'll see with Sarah and then we'll see with some other people where he's just trying to get under people's skin. That antagonistic approach is kind of fun to play around with and along with that comes some pretty intense scenarios. So the answer is Rudy.

Orphan Black has always been jam-packed full of action. What sort of physical preparations or training did you do to prepare for that challenge?

Ari Millen: I had to look like a military man! [Laughs]. It was good motivation to get off the couch, that's for sure. But other than that, I guess the diet. It's amazing what kind of willpower you can have when you're motivated that way. Unfortunately, it doesn't last but I was in the best shape of my life for the first little bit. [Laughs].

In addition to enjoying your work on Orphan Black, we have a memorable impression of you from Darknet. You seem to do really well with infusing this troublesome darkness into your work. What is it about this energy dynamic that you're drawn to playing?

Ari Millen: I don't know, I think bad guys are a lot more fun to play. But I don't know, I think I'm a nice guy! [Laughs]. They keep casting me this way but I'll take it!

What was your approach to playing all of these clones? Tatiana raved about your portrayal but she said your approach was different than hers.

Ari Millen: Certainly. Going back to the fact that they were self-aware, I guess the best way to describe it is, if you've ever met twins or triplets or, I don't know, I've never met quadruplets but if you don't know them that well, you could very easily confuse them until you get to know them and then you get to know their individual personalities. One of the set PAs on our show, he's one of triplets and I had also worked with his brother Anthony and mistaken his brother for him on a different show that I had done earlier in the year. It was a really good lesson. 'So, how do you find the differences within the similarities? What makes this guy uniquely himself?' That was the biggest challenge, because they were so similar. 'Mark's like this, so what motivates Seth to do something like this where in this scenario, Seth would do this but what would Rudy do or what would Miller do?' It was just comparing them against each other and finding those little things. Whereas with Project Leda, they grew up in different parts of the world so they can be as drastically different as possible and Tat can have free reign and explore and have fun with these huge differences. I would say that's the biggest difference between Castor and Leda, their upbringing.

Each season, Tatiana has played a core group of four to five clones and a few miscellaneous others. How many would you say are going to be in your core group that we'll see this season?

Ari Millen: [Laughs]. I feel like that might be a spoiler. We know of four, so I'll talk about four. Um, yeah, I'll say four.

Of the four, how many times did you have all four of them in the same scene or, what is the challenge of when there's four or even three in the same place? What's the toughest part when they all have to interact?

Ari Millen: Having two in a scene was hard enough so thankfully we didn't have much more than that. The difficult part of having multiples in a scene is just making sure that you're planning. All the acting training that I've ever had and most people would have is, you plan the scene as your character but if you have to play both sides of the scene, you've got to know both sets of motivations and plan out how they're going to execute. It's all about that first performance and the first half of shooting that one scene is to make sure that it's open enough for the other side of it to make sense. Don't set it in stone in the first couple ones otherwise you're stuck with it.

If you can tell us without it being too much of a spoilers, which one of your clones encountering Tatiana's clones is your favourite pairing?

Ari Millen: Well, coming into Orphan Black, first off my favourite clone was Helena. Mark got to meet her pretty quickly and it was mostly playing off of Sarah and Helena the whole second season and those two are my favourites. But in going into season 3, if I talked about anyone that would be a spoiler but I've already gotten to work with my favourites so I'm lucky.

Can you talk a bit about the audition process? What did you show them that convinced them that you could do this and take on this massive task?

Ari Millen: Oh man, if I knew that I would be pulling that all the time. That's probably better answered by John and Graeme but all I can say is I'm very thankful that they gave me the opportunity. I've had the best time of my life this season. I guess it made sense because of story, why Mark being a clone would be a good idea. But I wouldn't want to put words in their mouth. [Laughs]. I'm just happy they did.

What was it like stepping into this established world? Was everybody very welcoming to you?

Ari Millen: Yes, it's a dream set to be on. From craft to the writers room to all the actors that we work with, everyone is… and this is unique. This is not something I throw around lightly. Not every set that you go on is this cohesive and this motivated and supportive. It's a very wonderful group of people to be a part of and it makes it easier to take risks and be a clone. [Laughs]. That's no easy thing to tackle.

The show is nothing short of a phenomenon. What do you think it is that viewers connect so much with?

Ari Millen: The fact that there are clones, in one show or in one or I guess now two actors' performances, people can relate to different clones and different characters so it appeals to a broader audience. That's one aspect but it's also really smart writing. That's what attracted me initially. When I was watching, I really enjoyed the plot line. I just found it really clever. It's not too over-the-top that it lives in the world of your classic science-fiction. It's really rooted in reality with science that may very well exist at this point and we're just saying it does and this is the possibility of what could happen. It's giving strong voices to lots of talented actors and I think that's why it appeals to so many people.

What's not to love about it, right?

Ari Millen: Yes, exactly!

We've got a lot of American and Canadian accents going on. Any chance of getting more interesting accents going on?

Ari Millen: I don't know about that yet but I certainly tried my hardest this year. Thankfully, we have a dialect coach on set most of the time but I definitely tried to drop my Canadian-isms. These guys are supposed to be American so there are certain words that would come across as Canadian. They kept telling me, "Ari. Drop it. Drop the Canadian."

I know Tatiana has done this already but if you were given the chance, what would you think about playing a transgender Castor clone?

Ari Millen: You know, that's a very deep and serious question as far as the subject matter. I think what Tat and what the writers room did with Tony was no short of pretty incredible but that is something that is, it's not mainstream so much yet and if you're going to do it, you have to do it right. You have to do it proper justice. If that were ever something they would want to tackle with Castor, I would really hope that we would do it as best as possible and I would do as much research as possible. That is not something to throw around as a gimmick, that's real stuff. I think they did a beautiful job with Tony.

And are you hoping you get your own Clone dance party?

Ari Millen: [Laughs]. That would be cool!

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Read our MuseLed interview with Tatiana Malsany here

Orphan Black interviews from WonderCon 2015

Orphan Black season 3 premieres on Saturday, April 18, at 9PM on BBC America.

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